For the most up-to-date list, see my page on Google Scholar.


Draghi JA & Ogbunugafor, C. B. (2022). Exploring the expanse between theoretical questions and experimental approaches in the modern study of evolvability. Journal of Experimental Zoology Part B: Molecular and Developmental Evolution.


Bruger EL, Chubiz LM, Rojas Echenique JI, Renshaw CJ, Espericueta NV, Draghi JA, & Marx CJ (2021). Genetic context significantly influences the maintenance and evolution of degenerate pathways. Genome biology and evolution13(6), evab082.

Draghi, J. A. (2021). Asymmetric evolvability leads to specialization without trade-offs. The American Naturalist197(6), 644-657.

Svensson, E. I., Arnold, S. J., Bürger, R., Csilléry, K., Draghi, J., Henshaw, J. M., … & Runemark, A. Correlational selection in the age of genomics. Nature ecology & evolution 5, no. 5 (2021): 562-573.

Bruger, E. L., Chubiz, L. M., Rojas Echenique, J. I., Renshaw, C. J., Espericueta, N. V., Draghi, J. A., & Marx, C. J. (2021). Genetic context significantly influences the maintenance and evolution of degenerate pathways. Genome biology and evolution13(6), evab082.

Teufel, A. I., Liu, W., Draghi, J. A., Cameron, C. E., & Wilke, C. O. (2021). Modeling poliovirus replication dynamics from live time-lapse single-cell imaging data. Scientific reports11(1), 1-15.


Bono, L. M., Draghi, J. A., & Turner, P. E. (2020). Evolvability costs of niche expansion. Trends in Genetics36(1), 14-23.

Matthey‐Doret, R., Draghi, J. A., & Whitlock, M. C. (2020). Plasticity via feedback reduces the cost of developmental instability. Evolution Letters4(6), 570-580.


Ho, V., Herman-Bausier, P., Shaw, C., Conrad, K. A., Garcia-Sherman, M. C., Draghi, J., … & Rauceo, J. M. (2019). An Amyloid Core Sequence in the Major Candida albicans Adhesin Als1p Mediates Cell-Cell Adhesion. mBio10(5), e01766-19. [pdf]

Draghi, J. (2019). Phenotypic variability can promote the evolution of adaptive plasticity by reducing the stringency of natural selection. Journal of evolutionary biology. [pdf]

Draghi, J. (2019). Developmental noise and ecological opportunity across space can release constraints on the evolution of plasticity. Evolution & development, e12305. [pdf]

Zhao, L., Seth-Pasricha, M., Stemate, D., Crespo-Bellido, A., Gagnon, J., Draghi, J., & Duffy, S. (2019). Existing host range mutations constrain further emergence of RNA viruses. Journal of virology93(4), e01385-18. [pdf]

Draghi, J. (2019). Links between evolutionary processes and phenotypic robustness in microbes. In Seminars in cell & developmental biology (Vol. 88, pp. 46-53). Academic Press. [pdf]


Gilbert, K. J., Sharp, N. P., Angert, A. L., Conte, G. L., Draghi, J. A., Guillaume, F., Hargreaves, A. L., Matthey-Doret, R. & Whitlock, M. C. (2017) Local adaptation interacts with expansion load during range expansion: maladaptation reduces expansion load. The American Naturalist 189, no. 4 : 368-380. [pdf]

Draghi, J., & Whitlock, M. (2015). Robustness to noise in gene expression evolves despite epistatic constraints in a model of gene networks. Evolution69(9), 2345-2358. [pdf]

Draghi, J., & Whitlock, M. C. (2015). Overdominance interacts with linkage to determine the rate of adaptation to a new optimum. Journal of evolutionary biology28(1), 95-104. [pdf]

Schulte, M. B., Draghi, J. A., Plotkin, J. B., & Andino, R. (2015). Experimentally guided models reveal replication principles that shape the mutation distribution of RNA viruses. Elife4, e03753. [pdf]


Chou, H. H., Delaney, N. F., Draghi, J. A., & Marx, C. J. (2014). Mapping the fitness landscape of gene expression uncovers the cause of antagonism and sign epistasis between adaptive mutations. PLoS genetics10(2), e1004149. [pdf]

Draghi, J. A., & Plotkin, J. B. (2013). Selection biases the prevalence and type of epistasis along adaptive trajectories. Evolution67(11), 3120-3131. [pdf]

Draghi, J. A., & Whitlock, M. C. (2012). Phenotypic plasticity facilitates mutational variance, genetic variance, and evolvability along the major axis of environmental variation. Evolution66(9), 2891-2902. [pdf]

Turner, P. E., Draghi, J. A., & Wilpiszeski, R. (2012). High-throughput analysis of growth differences among phage strains. Journal of microbiological methods88(1), 117-121. [pdf]

Draghi, J. A., Parsons, T. L., & Plotkin, J. B. (2011). Epistasis increases the rate of conditionally neutral substitution in an adapting population. Genetics187(4), 1139-1152. [pdf]

Kryazhimskiy, S., Draghi, J. A., & Plotkin, J. B. (2011). In evolution, the sum is less than its parts. science332(6034), 1160-1161. [pdf]

Draghi, J. A., & Plotkin, J. B. (2011). Molecular evolution: Hidden diversity sparks adaptation. Nature474(7349), 45. [pdf]


Draghi, J. A., Parsons, T. L., Wagner, G. P., & Plotkin, J. B. (2010). Mutational robustness can facilitate adaptation. Nature463(7279), 353. [pdf]

Draghi, J., & Wagner, G. P. (2009). The evolutionary dynamics of evolvability in a gene network model. Journal of evolutionary biology22(3), 599-611. [pdf]

Draghi, J., & Wagner, G. P. (2008). Evolution of evolvability in a developmental model. Evolution: International Journal of Organic Evolution62(2), 301-315. [pdf]

Draghi, J. A., & Turner, P. E. (2006). DNA secretion and gene-level selection in bacteria. Microbiology152(9), 2683-2688. [pdf]